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Human remains found buried beneath a social services car park in Leicester are those of Richard III who was killed in battle in 1485, archaeologists confirmed today.
In an extraordinary discovery which rewrites the history books, the skeleton of the last of the Plantagenet kings was identified by DNA analysis after researchers traced his living descendants.
Investigators from the University of Leicester today revealed that the remains bore the marks of ten injuries inflicted shortly before his death.
More gruesome, however, was evidence of ‘humiliation’ injuries, including several head wounds - part of the skull was sliced away - a cut to the ribcage and a pelvic wound likely caused by an upward thrust of a weapon, through the buttock.
The hunchback king: The skeleton, unearthed in a dig last September, showed evidence of the same curvature of the spine and battle injuries thought to have been suffered by the last Plantagenet king
The face of a king: There were cheers from media who had gathered from around the world as the announcement was made at the University of Leicester this morning
The fatal blows? This image of the skull shows where Richard III was injured
This X-ray tomography image shows the two injuries which could have killed Richard: The area in the middle marked A is where the spine meets the skull. There are two injuries to the left (B) and right (C) of this that could have led to death if inflicted in life. The right hand injury, possibly from a halberd would have damaged the cerebellum. The left hand injury was probably caused by a sword and could also have been fatal
The skeleton was described of that of a slender male, in his late 20s or early 30s. Richard was 32 when he died.
Newly-released pictures also show a distinctive curvature of the spine synonymous with the hunchback king immortalised by Shakespeare.
There was, however, no evidence of a withered arm, which was also part of the Richard myth.
Speaking to 140 journalists who had travelled from across the world for the announcement, the university’s lead archaeologist Richard Buckley described the identity of the remains as ‘beyond reasonable doubt.'
‘It is the academic conclusion of the University of Leicester that the individual exhumed at Greyfriars in August 2012 is indeed King Richard III, the last Plantagenet King of England.’
The cut mark on the right rib of King Richard III: It is thought this and a number of other injuries found on the skeleton are evidence of 'humiliation injuries' inflicted after his death
Two vertebrae of king Richard III, showing some abnormal features relating to the scoliosis: The find corroborates historical accounts of Richard which described him as a hunchback
The blade wound to Richard's pelvis: This pelvic wound was likely caused by an upward thrust of a weapon through the buttock, researchers said
Another cut mark can be seen on the jaw bone of Richard III: Researchers identified ten wounds on the remains
Deputy registrar Richard Taylor described the discovery as ‘truly astonishing’ and said it could ‘prove to be one of the biggest archaeological discoveries of recent times’.
The long-awaited announcement was greeted by cheers.
The villain king: But there are those who suggests Richard III's bad reputation is more down to Tudor propaganda than his actual actions
Richard, depicted by William Shakespeare as a monstrous tyrant who murdered two princes in the Tower of London, died at the Battle of Bosworth Field, defeated by an army led by Henry Tudor.
According to historical records, his body was taken 15 miles to Leicester where it was displayed as proof of his death before being buried in the Franciscan friary.
The team from Leicester University set out to trace the site of the old church and its precincts, including the site where Richard was finally laid to rest.
They began excavating the city centre location in August last year and soon discovered the skeleton, which was found in good condition with its feet missing in a grave around 68cm (27in) below ground level.
It was lying in a rough cut grave with the hands crossed in a manner which indicated they were bound when he was buried.
To the naked eye, it was clear that the remains had a badly curved spine and trauma injuries to the rear of the head.
But archaeologists were keen to make no official announcement until the skeleton had been subjected to months of tests.
As they were found: The remains of King Richard III were found in a hastily dug grave beneath a council car park in Leicester last September, in what were once the precincts of Grey Friars church
The skull of the king as it was found by archaeologists: Trauma to the skeleton showed the king died after one of two significant wounds to the back of the skull - possibly caused by a sword and a halberd
Hunched in death as he was in life: The skeleton was found in good condition with its feet missing
The Battle of Bosworth: Richard, pictured on the white horse, was killed in battle more than 500 years ago at Bosworth field, in a battle which marked the end of his line and the rise of the Tudors
One woman's hunch led to the discovery of the skeleton which has now been proven to be that of Richard III.
Screenwriter Philippa Langley, pictured right, said she felt a chill on a hot summer's day as she walked through the area where it was thought he was buried.
The remarkable discovery of the remains, which, consistent with historical accounts of Richard, have both a curved spine back and wounded skull, was made last September.
Miss Langley was strolling across the car park used by Leicester social services while researching a play about the king when she felt a chill in August 2009.
'It was a hot summer and I had goosebumps so badly and I was freezing cold. I walked past a particular spot and absolutely knew I was walking on his grave,' she told the Sunday Times.
'I am a rational human being but the feeling I got was the same feeling I have had before when a truth is given to me.'
Miss Langley initially funded the excavation of what is now a Leicester City Council car park because she was '99 per cent certain' that the remains were those of Richard.
Miss Langley, who is a member of the Richard III Society, is working on a documentary charting the excavation for Channel 4 titled Richard III: The King in the Car Park, which has been made alongside the university academics and will be screened tonight.
She said the play that she began researching three years ago has been turned into a script for television and film, which is now 'getting serious interest from Los Angeles and in the UK'.
Speaking at today’s press conference, University of Leicester geneticist Dr Turi King described how researchers had traced Richard’s descendants to confirm the body was indeed that of England’s last medieval king.
These were Canadian born furniture maker Michael Ibsen, a direct descendant of the Richard’s sister Anne of York, and a second person who has asked to remain anonymous.
Dr King said: ‘The DNA sequence obtained from the Grey Friars skeletal remains was compared with the two maternal line relatives of Richard III.
‘We were very excited to find that there is a DNA match between the maternal DNA from the family of Richard III and the skeletal remains we found at the Grey Friars dig.’
The analysis showed the individual had a slender physique and severe scoliosis - a curvature of the spine - possibly with one shoulder visibly higher than the other.
This is consistent with descriptions of Richard III's appearance from the time, the researchers said today.
Trauma to the skeleton showed the king died after one of two significant wounds to the back of the skull - possibly caused by a sword and a halberd.
Dr Appleby said this was consistent with contemporary accounts of the monarch being killed after receiving a blow to the head.
The skeleton also showed a number of non-fatal injuries to the head and rib and to the pelvis, which is believed to have been caused by a wound through the right buttock.
Dr Appleby said these may have been so-called ‘humiliation injuries’ inflicted after his death.
‘The skeleton has a number of unusual features: its slender build, the scoliosis and the battle-related trauma,’ she said.
‘All of these are highly consistent with the information that we have about Richard III in life and about the circumstances of his death.
‘Taken as a whole, the skeletal evidence provides a highly convincing case for identification as Richard III.’
Skeletal analysis: Dr Jo Appleby presented the results of the analysis of the skeleton, which she said presented a 'highly convincing case' that it was Richard III
Confirmed 'beyond reasonable doubt': Lead researcher Richard Buckley for the first time shows the remains of King Richard III as they appeared in the grave found in the Grey Friars car park
Positive ID: Dr Turi King presents the findings of the DNA analysis which showed the skeleton did belong to King Richard III by matching them with Michael Ibsen, a descendant of Richard's maternal line
Maternal line descendant: Michael Ibsen provides the DNA sample which was used to prove the identity of the skeletal remains as those of Richard III
The bones had also undergone radiocarbon dating which indicated the man found had died sometime between 1485 and 1550 - consistent with historical records of the king’s death.
Archaeologists, historians and local tourism officials were all hoping for confirmation that the monarch's long-lost remains have been located.
So were the king's fans in the Richard III Society, set up to re-evaluate the reputation of a reviled monarch.
The search for the lost king: The announcement follows months of analysis of the remains since they were unearthed last September in a car park behind a council social services building in Leicester
We've been looking for you: Actors dressed as knights look where archaeologists found skeletal remains during an archaelogical dig to find the remains of King Richard III in Leicester
The spot in a Leicester car park where a set of remains were found which may be Richard III
Richard was immortalised in a play by William Shakespeare as a hunchbacked usurper who left a trail of bodies - including those of his two young nephews, murdered in the Tower of London - on his way to the throne.
Richard III remains an enigma - villain to many, hero to some. He ruled England between 1483 and 1485, during the decades-long tussle over the throne known as the Wars of the Roses.
His brief reign saw liberal reforms, including introduction of the right to bail and the lifting of restrictions on books and printing presses.
His rule was challenged, and he was defeated and killed at the Battle of Bosworth Field by the army of Henry Tudor, who took the throne as King Henry VII.
Richard III’s remains are expected to be re-interred in Leicester Cathedral.
New resting place: It is expected that Richard III's remains will now be re-interred at Leicester cathedral